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praises Alcmaeon, Callisthenes, and Herophilus for their work on the nature of the eye. He mentions that Alcmaeon excised an animal eye to study the optic nerve. However, there is no evidence that Alcmaeon himself dissected the eye or the skull. Based on this observation, and more rudimentary, Alcmaeon described the senses, except for the touch sense. These observations contributed to the study of medicine by establishing the connection between the brain and the sense

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