ribosomal DNA genotypes were considered together, it is argued that there are six ecotypes in the otherwise "mesic" classification.

Whole-plant studies also showed that the xeric and mesic types differed from each other for many characters such as the flag leaf, primary stem height, number of tillers, weight and number of seeds, dry weight, and flowering time, with the mesic ecotype being generally larger and more fecund, overall, than the xeric type; further, the flag leaves of the xeric type were