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showed in 1818 in his theorie d'analogue ("theory of homologues") that structures were shared between fishes, reptiles, birds and mammals. When Geoffroy went further and sought homologies between Georges Cuvier's embranchements, such as vertebrates and molluscs, his claims triggered the 1830 Cuvier-Geoffroy debate. Geoffroy stated the principle of connections, namely that what is important is the relative position of

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