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two β subunits, and two δ (sometimes γ: VnfG/AnfG) subunits. The delta subunits are homologous to each other, and the alpha and beta subunits themselves are homologous to the ones found in MoFe nitrogenase. The gene clusters are also homologous, and these subunits are interchangeable to some degree. All nitrogenases use a similar Fe-S core cluster, and the variations come in the cofactor metal. The δ/γ subunit helps bind the cofactor in the FeFe nitrogenase. Based on the timing of its evolution, the subunit in VFe

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