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appear during the Tremadocian. Tremadocian cephalopods were not very different from their Cambrian predecessors. Specimens of Ellesmeroceras and possibly Bassleroceras, found in Santa Rosita Formation, northwestern Argentina, show that cephalopods first migrated to the waters off western Gondwana already in the early Tremadocian. In the middle Tremadocian, cephalopods became more diverse

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