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<p>their <a href="page.php?w=symbionts">symbionts</a> to adequately <a href="page.php?w=photosynthesis">photosynthesize</a>, due to decreased light exposure.</p>

<p>The two main variables determining the <a href="page.php?w=geomorphology">geomorphology</a>, or shape, of coral reefs are the nature of the <a href="page.php?w=substrate_%28biology%29">substrate</a> on which they rest, and the history of the change in sea level relative to that substrate.</p>

<p>The approximately 20,000-year-old <a href="page.php?w=Great_Barrier_Reef">Great Barrier Reef</a></p><p>
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