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can sink to the ocean floor when radiolarians die and become preserved as part of the ocean sediment. These remains, as microfossils, provide valuable information about past oceanic conditions.

{\partial t} = \phi(\nabla^2) \mathbf{U} + G \mathbf{U}^2 - H\mathbf{U}V,

:

The function , taken to be the radius vector from the centre to any point on the surface of the membrane, was argued to be representable as a series of normalised Legendre functions.

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