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September 2013, Eurostar announced that its new service between London and Amsterdam, intended to begin operation in December 2016, would be operated by the trains. In April 2018 the scheduled service to Amsterdam started with two e320 trains per day. Due to remodelling of Amsterdam Central the direct reservice was paused from summer 2024 to February 2025, and thereafter its frequency and passenger capacity increased. Following Eurostar's merger with Thalys, the trains have also been used for routes in Mainland

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