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By the end of the 20th century, it was assumed that humans evolved from a chimpanzee-like ancestor. A 2015 study of Ardipithecus, an early human relative dated 4.4 mya, disputed this interpretation. The study concluded that humans, chimpanzees and bonobos evolved from a generalised species that lacked the derived traits of both genera. Subsequent studies disputed this, finding that the hand of Ardipithecus was adapted for suspension climbing and the heel

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