hypothesis, the archedictyon contained 6-8 longitudinal veins. These veins (and their branches) are named according to a system devised by John Comstock and George Needham--the Comstock-Needham system:

:Costa (C) - the leading edge of the wing

:Subcosta (Sc) - second longitudinal vein (behind the costa), typically unbranched

:Radius (R) - third longitudinal

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