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of which the hadrosaur body plan had little in the way of armor and speed.

This hypothesis led Ostrom to further conclude that ecology of hadrosaurs was more likely to be that of dry ground such as conifer forests, rather than swampy, aquatic environments, thought to be the case at the time. This idea was further justified by a 1922 paper that Ostrom rediscovered in 1964, which described the stomach contents of a mummified specimen of the hadrosaur Anatosaurus, which included conifer needles,

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