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with a partner of a different species (heterospecific mating) and losing fitness through hybridization. Inclusive fitness appears to be maximized in matings of intermediately related individuals.

Mechanisms

, five proposed mechanisms address the evolution of mate choice:
* Direct phenotypic benefits
* Sensory bias
* Fisherian runaway
* Indicator traits
* Genetic compatibility

Direct and/or indirect benefits drive the mating biases described

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