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b. 1221-1224.
** George Laskaris, brother of Theodore I Laskaris, b. 1211-1224?.
** Isaac Doukas Vatatzes, brother of John III Doukas Vatatzes, c. 1253- b. 1261.
** John Palaiologos (brother of Michael VIII), 1259-1260.
** Constantine Palaiologos (half-brother of Michael VIII),

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