and Nymphonidae (five pairs in Pentanymphon, six pairs in Sexanymphon).

Several alternatives had been proposed for the position homology of pycnogonid appendages, such as chelifores being protocerebral/homologous to the labrum (see text) or ovigers being duplicated palps. Conclusively, the classic, morphology-based one-by-one alignment to the prosomal appendages of other chelicerates was confirmed by both neuroanatomic