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a 2014 study concluded that these previous phylogenies, which were based on nucleic acid sequences, were subject to composition biases, and that, furthermore, phylogenies based on amino acid sequences suggested that the bryophytes are monophyletic after all. Since then, partially thanks to a proliferation of genomic and transcriptomic datasets, almost all phylogenetics studies based on nuclear and chloroplastic sequences have concluded that the bryophytes form a monophyletic group. Nevertheless, phylogenies based on mitochondrial sequences fail

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