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and A. sturio to form the most basal clade among sturgeons, and all other species being in a separate clade, with the various other species of Acipenser, Scaphirhynchus, Pseudoscaphirhynchus, and Huso to have varying levels of relationship with one another. The morphologically distinct "shovelnose" type sturgeons are deeply nested within the former "Acipenser" and "Huso" clades. The divergences within the group have also been found to be much deeper,

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