frequencies that occurs over time within a population.

Given the following:

# A particular locus on a chromosome and a given allele at that locus
# A population of N individuals with ploidy n, i.e. an individual carries n copies of each chromosome in their somatic cells (e.g. two chromosomes in the cells of diploid species)
# The allele exists in i chromosomes in the population

then the allele frequency is

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