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a simple complementation test. When the mutations in question are homozygous and recessive, complementation will ordinarily result in a normal (or "wild-type") phenotype if the mutations are in different genes (intergenic complementation). When the mutations are in different genes, each strain's genome supplies the wild-type allele to "complement" the mutated allele of the other strain's genome. Since the mutations are recessive, the offspring that is heterozygous mutant at each gene will display the wild-type

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