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spike-like cones, arranged in lax, cymes. Male spikes have collars closely arranged and ± hiding axis (less often somewhat laxly arranged), each collar with 20-80 flowers, often also with a whorl of sterile female flowers, apical whorl with sterile female flowers only. Female spikes are solitary or several in a panicle, often cauliflorous; involucral collars widely separated, each with 4-12 flowers.

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