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and non-reproductive groups. The division of labor creates specialized behavioral groups within an animal society, sometimes called castes. Eusociality is distinguished from all other social systems because individuals of at least one caste usually lose the ability to perform behaviors characteristic of individuals in another caste. (Narrow definitions, such as that of Crespi & Yagena, require that an irreversibly sterile caste be present, but this is not universally agreed on.) Eusocial colonies can be viewed as superorganisms.

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