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<p>, and <a href="page.php?w=Nonmanual_feature">nonmanual features</a>. Different sign languages make use of different handshapes.</p>

<p><big> Constraints </big></p>
<p>Possible handshapes are constrained by a variety of mechanic and neural factors. <a href="page.php?w=Evolution">Evolutionary</a> forces have led to some handshapes being easier or more natural for humans to produce than others. These tendencies can be summarized as follows:</p>

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# The selected (extended) finger is either the thumb or the index finger<br/>
# Neighboring fingers are coupled<br/></p><p>
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