can differentiate between a predator's shadow and changes in light caused by clouds. An evolutionary trade-off has led to a compromise between the eyes and the shell; as the size and complexity of the eyes increase, the mechanical performance of their shells decrease, and vice versa.

A relatively good fossil record of chiton shells exists, but ocelli are only present in those dating to or younger; this would make the ocelli, whose precise function is unclear, likely the most recent eyes to evolve.

Although chitons lack osphradia,

1.Previous
3.Next