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have demonstrated that intermediate-frequency sinusoidal patterns are optimally-detected by the retina due to the center-surround arrangement of neuronal receptive fields. In an intermediate spatial frequency, the peak (brighter bars) of the pattern is detected by the center of the receptive field, while the troughs (darker bars) are detected by the inhibitory periphery of the receptive field. For this reason, low- and high-spatial frequencies elicit excitatory and inhibitory impulses by overlapping frequency peaks and troughs in the center and

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