Pleistocene, from around 400,000-300,000 years ago onwards.

The woolly mammoth (Mammuthus primigenius) had emerged in Northeast Siberia from M. trogontherii by around 600-500,000 years ago, reaching the typical molar morphology of M. primigenius around 400,000 years ago. Mammoths with M. primigenius type molar morphology displaced those of M. trogontherii type in Europe over the course of the late Middle Pleistocene, which was largely complete by 200,000 years ago (~MIS