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as F. ariel. In 1914 the Australian ornithologist Gregory Mathews delineated five species, which remain valid. Analysis of ribosomal and mitochondrial DNA indicated that the five species had diverged from a common ancestor only recently--as little as 1.5 million years ago. There are two species pairs, the great and Christmas Island frigatebirds, and the magnificent and Ascension frigatebirds, while the fifth species, the

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