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there to be no morphological features that define the major clades within Isoetes, and no fossils are known that can be definitively assigned to the crown group.' While Wood et al. suggested a young origin dating to the early Cenozoic based on molecular clock estimates', the results were questioned by Wikström et al. (2023) who regarded the molecular clock as providing no firm evidence for the origin time of the genus, which could date to the Mesozoic

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