through calls, typically in the evening. Once a male has successfully attracted a mate, the process of amplexus begins, while the unsuccessful males are forced to continue their search for a mate through further calls.

The competition for a female mate among males is considered intense, and it is not uncommon for a male amphibian to attack an already-amplexed pair of amphibians. When a male amphibian attacks an amplexed pair of amphibians, he is trying to force the other male to release its grasp of the female, so he can then mate with