the main factors keeping ammonoid and brachiopod diversity low in this basin, with species of the genus Ceratites and two species of the nautiloid Germanonautilus being the only endemic cephalopods known from this basin, apart from occasional individuals of ammonoid species found in the Tethys that may have accidentally entered during stronger marine incursions into the basin.

Following the hyper saline Mittlerer Muschelkalk unit with its frequent evaporite deposits, a renewed opening of some of the gates to the Tethys brought full marine