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surrounds the mouth and is an upstream collecting system for suspension feeding. Its tentacles are hollow, with extensions of a coelomic space thought to be a mesocoel. The gut is U-shaped with the anterior mouth at the center of the lophophore. The anus, where present, is also anterior but is dorsal to the mouth. In the Bryozoa, it is outside the ring of the lophophore. The inarticulate brachiopods

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