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for vertebrates. Including one H⁺ for the transport reactions, this means that synthesis of one ATP requires 1 + 10/3 {{= 4.33}} protons in yeast and 1 + 8/3 {{= 3.67}} in vertebrates. This would imply that in human mitochondria the 10 protons from oxidizing NADH would produce 2.72 ATP (instead of 2.5) and the 6 protons from oxidizing succinate or ubiquinol would produce 1.64 ATP (instead of 1.5). This is consistent with experimental results within the margin of error

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