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in the taxonomy of poplars (Populus). Further areas of interest are cycads (Cycadales), conifers (Coniferales), morning glories (Convolvulaceae) and water hyacinth relatives (Pontederiaceae). His scientific work focuses on taxonomy, the formation of natural hybrids, paleobotany, and evolution, some of which led to significant changes in the taxonomy of conifers.

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