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on botany. He graduated in 1848 as doctor of philosophy with the thesis De forma et incremento stratorum crassiorum in plantarum cellula. He suggested that the plant cell wall growth occurred from the apposition from inside the cell not by adhesion from outside. He spent some time in London and Paris and joined the University of Berlin in 1851 as a privatdozent. His habilitation thesis was on the development of the oomycete of Saprolegnia ferax. In 1864 he succeeded Schlieden as professor of botany at the University of Jena.

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