as the cause of its spread--famously ending an outbreak by removing a public water pump handle in central London. His theory however only gained acceptance years after his death, through the discovery of the cholera bacterium.

Discovery and isolation

The discovery of the cholera toxin has widely been accredited to Robert Koch, a physician and microbiologist. In 1886, Koch hypothesised that Vibrio cholerae produced a substance that caused cholera symptoms.

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