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In the velvet belly lanternshark (Etmopterus spinax), a benthopelagic species, 17% of the bodyweight is liver of which 70% are lipids. Benthic rays and skates have smaller livers with lower concentrations of lipids; they are therefore denser than water and they do not swim continuously, intermittently resting on the seabed. Some fish have no buoyancy aids but use their pectoral fins which are so angled as to give lift

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