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<p>and P. fossilis. A 2022 study concluded that P. fossilis and P. spelaea represented a <a href="page.php?w=chronospecies">chronospecies</a> lineage, with most differences between the two species explainable by size differences.</p>

<p><big> Evolution </big></p>
<p>The lion lineage is thought to have originated in Africa, with fossils of the lineage on the continent extending back to the <a href="page.php?w=Late_Pliocene">Late Pliocene</a>, with the ancestors of Panthera fossilis migrating out of Africa during the <a href="page.php?w=Mid-Pleistocene_Transition">Early-Middle Pleistocene transition</a></p><p>
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