cones. The peduncles are glabrous. Each pollen cone has numerous spirally inserted microsporophylls each with two basal pollen sacs producing bisaccate pollen.

The female seed cones are solitary. Their peduncles may have small scale leaves. The cones consist of several sterile cone scales and one fertile cone scale with just one seed producing ovule. The sterile scales wither as the cone matures, unlike in the closely related genus Podocarpus where the scales fuse to form a fleshy receptacle. A part