take a year to become mature; they start out green or other bright colour, darkening as they develop to dark brown or black. The leaflike seed bracts are visible when young, and in some species remain so. The seeds sit in thin cups; each seed has a triangle-shaped wing. The male cones are pendulous (dangling) and resemble catkins; both pollen and seeds are wind-dispersed.

Evolution

Fossil history

The oldest pollen assignable to the genus dates to the

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