co-discovered genetic linkage through experiments with chickens and sweet peas.

In 1905 Punnett devised what is now called the Punnett square, a square diagram that is used to predict the genotypes of a particular cross or breeding experiment, described for the first time in the 2nd edition of his book.

In 1908, unable to explain how a dominant allele would not become fixed and ubiquitous in a population, Punnett introduced one of his problems to

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