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It is thought that it uses a di-heme active site, which is formed by the hemes of cytochromes b₅₉₅ and d. These two cytochromes are considered high-spin complexes, what is directly related to the electrons' spin. While other respiratory terminal oxidases which catalyze that same reaction have a heme-copper active site and use a proton pump, cytochrome bd has an active site with

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