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and archaea possess canonical SMC proteins that closely resemble their eukaryotic counterparts. These bacterial and archaeal SMCs form homodimers and associate with regulatory subunits to form condensin-like complexes, SMC-ScpAB. It is hypothesized that the eukaryotic ancestor (most likely the Asgard archaeon) possessed two types of SMC proteins: a canonical SMC (SMC^c) and a non-canonical SMC (SMC^nc). Gene duplications of these two ancestral

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