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teaching assignment was changed to molecular carcinogenesis. In 1995, together with Dinko Valerio, he created the PER.C6 cell line with enhanced traceability compared to the HEK 293 line, this time coming from retinal cells. The goal of Valerio was to use it for gene therapy but this was ultimately unsuccessful. In contrast to HEK 293 the PER.C6 cell line was patented by Leiden University and IntroGene (later Crucell)

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