non-African "Scalenodon" species. One Upper Manda cynodont, Aleodon, has also been found in the Dinodontosaurus assemblage zone of the Santa Maria Formation in Brazil.

Paleoenvironment

The Lifua Member sandstone was deposited by rivers. Meandering, probably sandy rivers were present. In the middle to upper Lifua bone accumulations, evidence suggests "that ponds and springs were a much more common occurrence on the Ruhuhu Basin floodplains at this

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