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anatomy. In 1903 Koltsov proposed that the shape of cells was determined by a network of tubules forming a skeleton which was later termed as the cytoskeleton. His studies were based on the sperm cells of various crustaceans and he examined their shapes and the physics of the structures. He saw the role of gel-sol transitions in the cytoplasm as key mechanisms for the cell structure. In 1927 Koltsov proposed that inherited traits would be inherited via a "giant hereditary

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