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genetic studies have suggested the choughs are basal to a group of Asian jay genera (Crypsirina, Dendrocitta, Platysmurus, Temnurus), or most recently, basal in the entire Corvidae. The genus Pyrrhocorax species differ from Corvus in that they have brightly coloured bills and feet, smooth, not scaled tarsi, and very short, dense nasal feathers. Choughs have

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