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that "Cleaning symbioses may not be mutualistic associations but rather one-sided exploitation. However, one must then ask why no counter-adaptation has evolved in clients to free them from this exploitation. If clients are the puppets of cleaners, then the fitness consequences of being exploited must be small". They quote as an example of an early position, C. Limbaugh writing in 1961: "From the standpoint of the philosopher of biology, the extent of cleaning behavior in the ocean emphasizes the role of co-operation in nature as opposed to the

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