living on the remote island of Skokholm. These small seabirds make one of the longest migrations of any bird--10,000 kilometres--but return to the exact nesting burrow on Skokholm year after year. This behaviour led to the question of how they navigated.

Mechanisms

Lockley began his book Animal Navigation with the words:

Many mechanisms of spatial cognition have been proposed for animal navigation: there is evidence for a number of them. Investigators

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