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<p>have evolved eight times, four of these being organs powerful enough to deliver an electric shock. Each such group is a <a href="page.php?w=clade">clade</a>. Most electric organs evolved from myogenic tissue (which forms muscle), however, one group of <a href="page.php?w=Gymnotiformes">Gymnotiformes</a>, the <a href="page.php?w=Ghost_knifefish">Apteronotidae</a>, derived their electric organ from neurogenic tissue (which forms nerves). In Gymnarchus niloticus (the African knifefish), the tail, trunk, hypobranchial, and eye muscles are incorporated</p><p>
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