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Archaeosporales), due to the similarity of Horodyskia's bead-like structures to the bladders of early growth stage Geosiphons. Horodyskia is almost certainly a eukaryote due to its large size and inferred cell wall, although likely a quite simple protist since its "beads" are quite cell-like in appearance. Its exact placement is unclear, as although a giant-celled alga is the best match, modern examples of these do not form lines of cells in a gelatinous matrix, alongside having rhizoids unlike Horodyskia.

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