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molecule" which would be made up of "two mirror strands that would replicate in a semi-conservative fashion using each strand as a template". Koltsov used the expression omnis molecula ex molecula (every molecule comes from another molecule) based on Virchow's idea that all cells came from other cells. Around 1922, he began to conduct mutation experiments, subjecting Drosophila and crustaceans to X-ray radiation, some of which were influenced by discussions with H. J. Muller. He suggested that mutagenesis

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