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(B. verrucosissimus) and the Japanese common toad (B. japonicus). The European common toad (Bufo bufo) seems to have arisen more recently. It is believed that the range of the ancestral form extended into Asia but that isolation between the eastern and western species complexes occurred as a result of the development of the Central Asian Deserts during the Middle Miocene. The exact taxonomic relationships between these species remains unclear. A serological

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